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Summary of succession on sand dunes
The vegetation succession
in sand dunes is often described in a sequence from open mobile dunes to more
stable types as the growing vegetation traps sand. This is true for sand dunes
in temperate regions, but not those of drier desert lands which are not the
subject of this guidance. The zonation which develops reflects the physical
characteristics affecting sand transport rates and deposition acting in concert
with the vegetation and other biological factors (many references describe this
but see Packham & Willis 1997 pp. 153-169).
The development of strandline
depends on the presence of debris and plants on the beach which help trap sand.
Foredune
development in north west Europe is directly related to the ability of plants
such as Elytrigia juncea and Ammophila
arenaria to withstand burial by sand together with the other stresses,
such as water availability, in what is an inhospitable environment. [Similar
species occupy the same niche in America, where Ammophila
breviligulata replaces the ubiquitous Ammophila
arenaria of Europe.]
Once the main body of the dune is formed other processes come into play and
initiate a sequence from mobile foredunes
and 'yellow dunes' to 'grey' semi-fixed dunes with grassland, heath, scrub and
woodland. A typical
sequence can be established which helps form the basis for considering management
and restoration. The original calcium content of the sand, and the age of the
dune soil (including the degree of leaching) helps to determine whether succession
is to calcareous dune
grassland or dune heath. This sequence rarely occurs in a straightforward
progression from one stage to the next as depicted. Blow-outs
occur with or without the intervention of man and can be the precursors of dune
slacks. Similarly the re-profiling of dune ridges under the influence
of changing wind patterns brings an infinitely variable topography, the origins
of which may be difficult or impossible to unravel. The complex topography,
spatial and temporal variation helps to sustain a particularly diverse flora
which includes many rare species and fauna.
References
Packham, J.R. & Willis, A.J.
1997.
Ecology of Dunes, Salt Marsh and Shingle. Chapman & Hall, London, 335 pp.
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